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Official statistic on seabird breeding population changes

This page hosts the Official Statistic ‘Seabird population changes in the UK: 1986–2024’, published on 19 March 2026.

Official Statistic description

  • This publication reports changes in UK seabird breeding abundance from 1986 to 2024 and productivity from 2023 to 2024. It is produced from data gathered by the Seabird Monitoring Programme (SMP), which conducts whole colony and plot-scale surveys at coastal and inland sites. In 2024, 551 UK sites were surveyed for abundance and 129 for productivity.
  • Changes in abundance, productivity, or both are presented for 22 of the 25 regularly breeding UK seabird species. However, monitoring of European Storm‑petrel, Leach’s Storm‑petrel and Mediterranean Gull is too sparse or unrepresentative to produce reliable figures.
  • Figures for country level species-specific changes in abundance or productivity are produced where sufficient data are available.
  • No figures are presented for Northern Ireland in 2024 due to insufficient data.
  • A summary of the survey and analytical methods used is presented in Section 5.

The SMP is a partnership jointly funded by the Joint Nature Conservation Committee (JNCC) and British Trust for Ornithology (BTO), in association with the Royal Society for the Protection of Birds (RSPB), with fieldwork conducted by both volunteer and professional surveyors. The programme is supported by a network of organisations that form an Advisory Group and by the SMP Steering Group. Steering Group meetings are also attended by the Statutory Nature Conservation Bodies: Department of Agriculture, Environment and Rural Affairs, Northern Ireland (DAERA), Natural Resources Wales (NRW), Natural England (NE) and NatureScot.

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Scope of Statistics

The statistic presents results for the UK and for England, Scotland and Wales. UK figures include data collected by the SMP from Northern Ireland, but not from the Channel Islands or the Isle of Man.

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Results

Summary tables showing abundance changes and productivity estimates for those seabird species with sufficient data are presented in Sections 3.2–3.5. SMP breeding abundance figures are expressed as percentage change over two timeframes: long‑term change (LT) and 24‑year change. Unless otherwise noted, the LT change figure spans the duration of the SMP (1986–2024), while the 24‑year change figure covers 2000–2024, with 2000 selected as it is the midpoint of the Seabird 2000 national breeding seabird census (1998–2002) (Mitchell et al. 2004).

The method for calculating breeding abundance change is described in Section 5. Increases or declines in abundance for a particular species have been considered to be significant over the long term when the 95% confidence intervals of the 2024 estimate do not overlap the 1986 baseline index of 100. Similarly, change is considered significant over the 24-year period when the 95% confidence intervals of the estimate for 2024 do not overlap with the index value for 2000. These results are marked with an asterisk in Tables 1–4. Confidence intervals are not currently available for the productivity estimates (see JNCC 2014 for details), so it is not possible to assess statistical significance or provide a measure of uncertainty for these values.

Additional detail is available from the BTO Bird Trends Explorer where species-specific breeding seabird abundance and breeding performance (productivity) graphs from 1986 to 2024 can be produced.

The SMP annual report (Harris et al. 2026) includes the results presented here within species-specific accounts. These also provide information that sits outside of the SMP Official Statistic on distribution, diet and breeding behaviour, and outline key threats and conservation actions.

 

Summary of results

The results presented in Tables 1–4 show that, in 2024, the number of significant declines for seabird species where sufficient data are available, both at a UK and country level, largely outnumber those showing a significant increase.

For the UK, abundance change figures for a total of 17 breeding seabird species could be estimated (Table 1), and a range of species showed significant declines in abundance over both the long-term (1986–2024) and 24-year (2000–2024) periods. Significant declines were apparent for Fulmar, Shag, Lesser Black-backed Gull (natural nesters only), Herring Gull (natural nesters only), Kittiwake, Sandwich Tern and Common Tern over both periods, while the decline for Arctic Skua was significant across the long-term period but not the 24-year period. Razorbill was the only species to show a significant increase (across the long-term time period only).

In England, abundance change figures could be produced for nine species (Table 2), and of these only Common Tern showed a significant change, with a decline in abundance over both the long-term (1986–2024) and 24-year (2000–2024) periods.

In Scotland, abundance change could be estimated for 13 species (Table 3), with many showing a similar pattern of decline to the UK, although there were some differences in the species affected and the extent of the effects. Significant decreases were shown for Fulmar, Shag, Herring Gull (natural nesters only), Great Black-backed Gull, Kittiwake, and Common Tern across both the long-term (1986–2024) and 24-year (2000–2024) periods. There were significant declines for Arctic Skua and Little Tern across the long-term period, but not for the 24-year period. No significant increases were identified.

In Wales, abundance change figures could be produced for eight species (Table 4). Razorbill showed significant increases over both time periods, whilst Great Black-backed Gull increased significantly over the long-term period (1986–2024) but not over the 24-year period (2000–2024). Kittiwake abundance declined significantly over both periods.

Changes in productivity estimates between 2023 and 2024 varied greatly between species and regions (Tables 1–4). Notable results included Sandwich Tern productivity estimates increasing across the UK, England and Scotland, whereas Shag productivity estimates declined in the UK, Scotland and Wales. Arctic Skua had a particularly poor year for productivity in 2024, with an estimate of only 0.03 chicks/fledged per pair in Scotland.

Possible drivers of change for the observed results are described in Section 4.1.

UK: Abundance and productivity changes for 22 species

  • In the UK, Razorbill was the only breeding seabird species to show a significant long‑term (1986–2024) increase in abundance (+110%) (Table 1).
  • Eight species declined significantly in abundance over the same period, with the steepest declines observed in Arctic Skua (−93%), Lesser Black‑backed Gull (natural nesters, −72%) and Shag (−71%) populations.
  • Over the 24‑year period (2000–2024), no species increased significantly. The strongest significant declines were observed in Lesser Black‑backed Gull (natural nesters, −81%), Shag (−65%) and Common Tern (–60%) populations.
  • From 2023 to 2024, the largest increases in productivity estimates (chicks fledged per pair) were recorded for Black‑headed Gull (0.30 to 0.72), Sandwich Tern (0.17 to 0.58) and Great Black‑backed Gull (1.04 to 1.24).
  • Over the same period, the largest decreases in productivity estimates were observed for Arctic Skua (0.58 to 0.03), Shag (1.31 to 0.93) and Little Tern (0.61 to 0.25).
Table 1: UK SMP breeding abundance change and productivity. (Select the arrows icon for an expanded view)

Species and count unit (see Section 5 for unit definitions)

Breeding Abundance
Change %

Productivity
(chicks fledged per pair)

Sites 2024

LT change (1986-2024)

24-yr change (2000-2024)

2023

Sites

2024

Sites

Fulmar (AOS)

173

-35*

-32*

0.33

35

0.29

35

Manx Shearwater

-

-

-

0.60

3

0.60

1

Gannet

-

-

-

0.60

6

0.53

7

Cormorant (AON)

49

-8

-16

-

-

-

-

Shag (AON)

116

-71*

-65*

1.31

20

0.93

17

Arctic Skua (AOT)

11

-93*

-89

0.58

2

0.03

4

Great Skua

-

-

-

0.44

4

0.17

6

Black-headed Gull (AON)

75

Coastal nesters°

-18

Coastal nesters°

-37

Coastal nesters°

0.30

All nesters

29

All nesters

0.72

All nesters

30

All nesters

Common Gull
All nesters

-

-

-

0.62

11

0.43

11

Lesser Black-backed Gull (AON) 
Natural nesters^

76

-72*

-81*

0.48

13

0.40

11

Herring Gull (AON)
Natural nesters^

169

-51*

-47*

0.52

15

0.69

19

Great Black-backed Gull (AON)

98

-16

-20

1.04

11

1.24

9

Kittiwake (AON)

99

-55*

-38*

0.78

28

0.72

30

Sandwich Tern (AON)

15

-26*

-24*

0.17

6

0.58

11

Roseate Tern (AON)

3

-61

125

-

-

0.85

2

Common Tern (AON)

103

-63*

-60*

0.43

29

0.27

37

Arctic Tern (AON)

52

-25

-29

0.11

16

0.11

19

Little Tern (AON)

37

-19

-2

0.61

44

0.25

20

Guillemot (IND)

105

40

18

0.46

11

0.51

8

Razorbill (IND)

123

110*

52

0.55

12

0.44

9

Black Guillemot (IND)

46

3

61

-

-

-

-

Puffin

-

-

-

0.49

6

0.56

8

Table 1 footnotes:

* Significant abundance changes (where 95% confidence intervals, which reflect imputation uncertainty (see Sections 4 and 5 for details), do not overlap with the 1986 baseline for the long-term change, or the 2000 index value for the 24-year change).

^ Natural nesters defined as breeding on moors, cliffs, marshes, beaches and other areas of natural or semi-natural habitat.

° Coastal nesters defined as colonies located within 5 km of the mean high water mark.

 

England: Abundance and productivity changes for 10 species

  • In England, the only seabird species to show a significant change in breeding abundance was Common Tern, which declined by 47% and 51% over the long‑term (1986–2024) and 24‑year (2000–2024) periods, respectively (Table 2).
  • From 2023 to 2024, the largest increases in productivity estimates (chicks fledged per pair) were recorded for Sandwich Tern (0.20 to 0.77) and Arctic Tern (0.19 to 0.41).
  • Over the same period, the largest decreases in productivity estimates were observed for Common Tern (0.90 to 0.38) and Little Tern (0.76 to 0.26).
Table 2: England SMP breeding abundance change and productivity. (Select the arrows icon for an expanded view)

Species and count unit (see Section 5 for unit definitions)

Breeding Abundance
Change %

Productivity
(chicks fledged per pair)

Sites 2024

LT change (1986-2024)

24-yr change (2000-2024)

2023

Sites

2024

Sites

Fulmar (AOS)

34

-40

-36

0.47

10

0.41

12

Cormorant (AON)

22

26

-8

-

-

-

-

Black-headed Gull (AON)
Coastal nesters°

52

-5

-34

-

-

-

-

Herring Gull (AON) 
Natural nesters^

59

-77

-73

-

-

-

-

Kittiwake (AON)

22

-43

-26

0.76

8

0.67

9

Sandwich Tern (AON)

9

-27

-22

0.20

4

0.77

5

Common Tern (AON)

60

-47*

-51*

0.90

21

0.38

22

Arctic Tern (AON)

9

-50

-38

0.19

4

0.41

6

Little Tern (AON)

20

-15

4

0.76

35

0.26

16

Guillemot

-

-

-

0.59

2

0.60

2

Table 2 footnotes:

* Significant abundance changes (where 95% confidence intervals, which reflect imputation uncertainty (see Sections 4 and 5 for details), do not overlap with the 1986 baseline for the long-term change, or the 2000 index value for the 24-year change).

^ Natural nesters defined as breeding on moors, cliffs, marshes, beaches and other areas of natural or semi-natural habitat.

° Coastal nesters defined as colonies located within 5 km of the mean high water mark.

 

Scotland: Abundance and productivity changes for 18 species

  • In Scotland, no breeding seabird species showed a significant long‑term (1986–2024) increase in abundance (Table 3).
  • Over the same period, eight species declined significantly, with the steepest long‑term declines observed in Arctic Skua (−93%), Little Tern (−75%), and Common Tern (−73%) populations.
  • Over the 24‑year period (2000–2024) the pattern is similar, with no significant increases and the strongest significant declines in Common Tern (−67%), Shag (−62%) and Great Black-backed Gull (–57%) populations.
  • From 2023 to 2024, the largest increases in productivity estimates (chicks fledged per pair) were recorded for Great Black‑backed Gull (1.00 to 1.23), Sandwich Tern (0.11 to 0.26), and Puffin (0.48 to 0.58).
  • Over the same period, the largest decreases in productivity estimates were observed for Arctic Skua (0.58 to 0.03), Shag (1.26 to 0.83), and Great Skua (0.44 to 0.17).
Table 3: Scotland SMP breeding abundance change and productivity.(Select the arrows icon for an expanded view)

Species and count unit (see Section 5 for unit definitions)

Breeding Abundance
Change %

Productivity
(chicks fledged per pair)

Sites 2024

LT change (1986-2024)

24-yr change (2000-2024)

2023

Sites

2024

Sites

Fulmar (AOS)

102

-36*

-32*

0.35

12

0.29

15

Gannet

-

-

-

0.59

4

0.64

5

Shag (AON)

74

-70*

-62*

1.26

13

0.83

10

Arctic Skua (AOT)

11

-93*

-89

0.58

2

0.03

4

Great Skua

-

-

-

0.44

4

0.17

6

Black-headed Gull
All nesters

-

-

-

0.49

5

0.49

4

Lesser Black-backed Gull (AON)
Natural nesters^

15

-42

-46

-

-

-

-

Herring Gull (AON)
Natural nesters^

75

-54*

-43*

-

-

-

-

Great Black-backed Gull (AON)

62

-59*

-57*

1.00

3

1.23

5

Kittiwake (AON)

63

-53*

-41*

0.71

14

0.71

15

Sandwich Tern

-

-

-

0.11

1

0.26

1

Common Tern (AON)

27

-73*

-67*

0.45

4

0.26

7

Arctic Tern (AON)

34

-35

-35

0.18

9

0.10

9

Little Tern (AON)

16

-75*

-59

0.50

7

0.41

4

Guillemot (IND)

70

-29

-38

0.46

6

0.55

5

Razorbill (IND)

78

63

4

0.53

6

0.51

3

Black Guillemot (IND)

11

-44

-14

-

-

-

-

Puffin

-

-

-

0.48

4

0.58

4

Table 3 footnotes:

* Significant abundance changes (where 95% confidence intervals, which reflect imputation uncertainty (see Sections 4 and 5 for details), do not overlap with the 1986 baseline for the long-term change, or the 2000 index value for the 24-year change).

^ Natural nesters defined as breeding on moors, cliffs, marshes, beaches and other areas of natural or semi-natural habitat.

 

Wales: Abundance and productivity changes for 8 species

  • In Wales, significant long‑term (1986–2024) increases in abundance were observed in Razorbill (+256%) and Great Black-backed Gull (+170%) populations (Table 4).
  • Over the same period, a significant decline was observed in the Kittiwake (−56%) population.
  • Over the 24‑year period (2000–2024), Razorbill abundance increased significantly (+141%) and the only significant decline was in the Kittiwake population (-53%).
  • From 2023 to 2024, the largest increase in productivity estimates (chicks fledged per pair) was recorded for Herring Gull (0.55 to 0.81).
  • Over the same period, the only decrease in productivity estimates was observed for Shag (1.63 to 1.59)
Table 4: Wales SMP breeding abundance change and productivity. (Select the arrows icon for an expanded view)

Species and count unit (see Section 5 for unit definitions)

Breeding Abundance
Change %

Productivity
(chicks fledged per pair)

Sites 2024

LT change (1986-2024)

24-yr change (2000-2024)

2023

Sites

2024

Sites

Fulmar (AOS)

23

-4

-26

0.34

4

0.49

1

Cormorant (AON)

8

-29

-7

-

-

-

-

Shag (AON)

16

-7

-1

1.63

4

1.59

2

Lesser Black-backed Gull (AON)
Natural nesters^

17

-76

-79

-

-

-

-

Herring Gull (AON)
Natural nesters^

29

-18

-31

0.55

4

0.81

4

Great Black-backed Gull (AON)

12

170*

76

-

-

-

-

Kittiwake (AON)

10

-56*

-53*

0.47

2

0.61

2

Razorbill (IND)

30

256*

141*

-

-

-

-

Table 4 footnotes:

* Significant abundance changes (where 95% confidence intervals, which reflect imputation uncertainty (see Sections 4 and 5 for details), do not overlap with the 1986 baseline for the long-term change, or the 2000 index value for the 24-year change).

^ Natural nesters defined as breeding on moors, cliffs, marshes, beaches and other areas of natural or semi-natural habitat.

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Interpretation notes

Drivers of change

  • From 2021 onwards, High Pathogenicity Avian Influenza (HPAI) caused major mortality and rapid population decline in several UK seabird species, most notably Great Skua and Gannet (Tremlett et al. 2024). Although HPAI outbreaks were less prevalent in 2024, they may continue to affect survival and productivity in future years.
  • Climate‑related changes, including warming seas, have caused shifts in the distribution, timing and availability of key prey species, leading to reduced foraging efficiency and breeding success in many seabird species. More frequent extreme weather events, also linked to climate change, such as winter storms and prolonged wet periods during the breeding season, can reduce overwinter adult survival, cause nest failures and increase chick mortality.
  • Pressures from non‑native predators, such as rats and American Mink, or in some cases from native predators, can lead to the loss of eggs, chicks and adults.
  • Fisheries interactions, including seabird bycatch in gillnets and on longlines, can reduce adult survival, while reductions in fishery discards have affected the food supply for species that scavenge from fishing boats. Reduced access to discards may force birds to forage further or switch to lower‑quality prey, potentially reducing adult condition and breeding success.
  • Human disturbance, including recreational activity near colonies and egg collection, can reduce breeding success in sensitive species, particularly among terns and gulls.
  • Renewable energy infrastructure, especially offshore wind farms, may displace some species from preferred foraging areas, increasing energetic costs, or create collision risks for species that fly at rotor height.
  • Habitat change, including erosion, vegetation change and loss of suitable nesting areas, may limit breeding opportunities for species such as Little Tern and Mediterranean Gull.

For a detailed review of UK breeding seabird population drivers of change, see Seabirds Count: A census of breeding seabirds in Britain and Ireland (2015–2021) (Burnell et al. 2023).

Confidence in results and caveats

  • SMP surveys and analytical approaches follow standardised peer-reviewed methods to ensure results are comparable between survey sites and over time. All data undergo a combination of automated and manual validation and verification. See Walsh et al. (1995), JNCC (2014) and Harris et al. (2026) for full details.
  • Valid abundance and productivity figures require sufficient spatial and temporal coverage, and species with challenging survey requirements (e.g. burrow‑nesters, nocturnal or remote breeders) are often under-represented. Results for these groups have lower confidence or cannot be produced, for example where fewer than the required proportion of colonies are monitored (see Section 5 for criteria), leading to gaps in some species and regions.
  • Abundance index graphs showing 95% confidence intervals are presented in the SMP annual report (Harris et al. 2026) and on the BTO Bird Trends Explorer.
  • Results for species with sparse datasets carry wider confidence intervals and should be interpreted cautiously.
  • Assessment of the significance of abundance changes considers whether the confidence intervals of the estimate for 2024 overlap the index values for 1986 (long-term change) or 2000 (24-year change). These assessments reflect the relative level of sampling effort in the relevant years.
  • Imputation is used to estimate missing annual counts, with uncertainty quantified through bootstrapping (see Section 5). Confidence intervals only reflect uncertainty in the imputation of missing data. They do not reflect uncertainty in abundance counts and, therefore, the overall uncertainty in the population estimates, so caution should be applied when interpreting the significance of reported population change figures. Due to sparse datasets for several species, especially at a country level, there is a high risk of false negatives when assessing the significance of population changes.
  • The indices used to assess population change are unsmoothed and therefore percentage changes could be influenced by extreme annual estimates.
  • Productivity estimates rely on Generalised Linear Mixed Models, but the absence of confidence intervals for productivity values limits the ability to assess statistical uncertainty in annual estimates.
  • COVID‑19‑related disruptions in 2020 and low sample sizes in some later years created gaps in productivity and abundance data, meaning estimates for these years are either interpolated or excluded from model fitting.
  • Some species’ datasets are dominated by a small number of long‑running study sites, particularly in Scotland, which improves consistency but may reduce the representativeness of UK‑wide figures where regional differences exist.
  • Changes in monitoring effort during national census years or periods of targeted surveying can temporarily increase coverage, but these peaks may create inconsistencies when compared with typical annual effort.
  • For several gull species, inland or urban colonies are not well covered because of survey difficulty, meaning reported abundance change figures reflect only natural or coastal nesters and should not be assumed to represent entire populations. Natural nesters are defined as breeding on moors, cliffs, marshes, beaches and other areas of natural or semi-natural habitat, whilst coastal nesters are defined as colonies located within 5 km of the mean high water mark.
  • A 2024 review of the SMP sampling strategy (O’Hanlon et al. 2024) found that some abundance and productivity trends were imprecise, absent, or geographically limited. Its recommendations included expanding monitoring in under-represented regions and inland areas, covering more species and colony sizes and revising trend analysis methods. Measures to implement these improvements are planned but will take time to complete.

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Methods

  • Annual SMP monitoring focuses on breeding abundance and productivity, collected through whole‑colony counts or fixed plots.
  • Standardised monitoring methods are encouraged, as recommended in the Seabird Monitoring Handbook (Walsh et al. 1995), to allow colonies, plots and nest sites to be compared consistently across species and years.
  • Breeding abundance data are collected using standard units such as Apparently Occupied Nest (AON), Apparently Occupied Site (AOS), Apparently Occupied Territory (AOT) or Individual (IND) counts of adults, depending on the species, with survey timing also guided by species‑specific recommendations in the Seabird Monitoring Handbook (Walsh et al. 1995).
  • Productivity is recorded as the number of chicks fledged per breeding pair, typically using repeat visits across the season at representative plots within each colony.
  • Analytical methods are described in Methods of analysis for production of indices of abundance and estimation of productivity (JNCC 2014) and in the ‘Background and methods’ section of the SMP annual report (Harris et al. 2026).
  • Percentage changes in breeding abundance are calculated by comparison of the annual abundance estimate for 2024 with those for 1986 (long-term change) or 2000 (24-year change) using the following formula:

Abundance change % = (Final index value - Initial index value) x 100
                                                       Initial index value

where the Final index value is that for 2024 and the Initial index value is that for 1986 for the long-term change and 2000 for the 24-year change.

  • The abundance trend analysis, which produces the annual indices used to calculate the long-term and 24-year abundance change values, requires at least three years of data per site across the monitoring period; species‑level trends are only produced when at least 15% of the underlying SMP data contains actual rather than imputed counts across the period assessed.
  • Missing annual counts are estimated through an imputation method that uses weighted mean values from non-missing counts across years, with uncertainty quantified by bootstrapping to generate confidence intervals.
  • Productivity estimates are generated using Generalised Linear Mixed Models, incorporating binomial or Poisson error structures and site‑level random effects; confidence intervals are not currently available for these.
  • Four Key Sites (Canna, Fair Isle, Isle of May and Skomer Island) provide additional demographic data, including diet, phenology and adult survival, to support interpretation of observed changes.
  • Additional triennial surveys at St Kilda, Orkney and Bullers of Buchan strengthen coverage for species or regions where annual sampling is limited.
  • National censuses, such as Seabird 2000 (1998–2002) (Mitchell et al. 2004) and Seabirds Count (2015–2021) (Burnell et al. 2023), provide population baselines which are used to contextualise annual SMP figures and assess long‑term abundance change.
  • All data are submitted through SMP Online, validated by BTO before analysis, and compiled into annual Official Statistics.

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Involvement and contacts

  • This statistic was produced by the SMP partnership, with BTO having primary responsibility.
  • The Lead Statistician was Nina O’Hanlon, Senior Research Ecologist at BTO (smp@bto.org).
  • Quality assurance was undertaken by BTO, JNCC and RSPB.
  • The SMP is delivered jointly by BTO and JNCC, in association with RSPB, and supported by an advisory network spanning government agencies, NGOs and local partners.
  • A dedicated BTO programme coordinator acts as the main point of contact for survey support, participation enquiries and coordination.
  • The data are published as a JNCC Official Statistic. If you have any queries, please contact us.

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Relation to other National and Official Statistics

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References

Burnell, D., Perkins, A.J., Newton, S.F., Bolton, M., Tierney, T.D. & Dunn, T.E. (eds.) (2023) Seabirds Count: a census of breeding seabirds in Britain and Ireland (2015–2021). Lynx Nature Books, Barcelona

Harris, S.J., Baker, H., Balmer, D.E., Bolton, M., Burton, N.H.K., Clarke, J.A.E., Dunn, T.E., Evans, T.J., Hereward, H.F.R., Humphreys, E.M., Langlois Lopez, S., Money, S., Taylor, E. A. and O’Hanlon, N.J. (2026) Seabird Population Trends and Causes of Change: 1986–2024, the annual report of the Seabird Monitoring Programme. BTO Research Report 795. British Trust for Ornithology, Thetford.

Joint Nature Conservation Committee (JNCC) (2014) Methods of analysis for production of indices of abundance and estimation of productivity. JNCC. Available at: https://data.jncc.gov.uk/data/701c338f-ed54-43da-a61c-254cb79698b8/Analysis-methods.pdf

Mitchell, I., Newton, S.F., Ratcliffe, N. & Dunn, T.E. (eds.) (2004) Seabird populations of Britain and Ireland: results of the Seabird 2000 census (1998-2002). T. & A.D. Poyser, London.

O’Hanlon, N.J., Harris, S.J., Thaxter, C.B., Boersch‑Supan, P.H., Robinson, R.A., Balmer, D.E. & Burton, N.H.K. (2024) Seabird population and demographic monitoring in the UK: a review and recommendations for future sampling. BTO Research Report 754. British Trust for Ornithology, Thetford. ISBN 978‑1‑912642‑49‑6. Available at https://www.bto.org/our-work/science/publications/reports/bto-research-reports/00754-seabird-population-and-demographic-monitoring-uk-review

Tremlett, C.J., Morley, N. & Wilson, L.J. (2024) UK seabird colony counts in 2023 following the 2021–22 outbreak of Highly Pathogenic Avian Influenza. RSPB Research Report No. 76. Sandy, Bedfordshire: RSPB Centre for Conservation Science, RSPB. ISBN 978‑1‑905601‑72‑1.

Walsh, P.M., Halley, D.J., Harris, M.P., del Nevo, A., Sim, I.M.W. & Tasker, M.L. (1995) Seabird Monitoring Handbook for Britain and Ireland. JNCC/RSPB/ITE/Seabird Group, Peterborough.

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